Colour banding

Colour banding

Colour banding is a subtle form of posterisation in digital images, caused by the colour of each pixel being rounded to the nearest of the digital colour levels. While posterisation is often done for artistic effect, colour banding is an undesired artefact. In 24-bit colour modes, 8 bits per channel is usually considered sufficient to render images in Rec. 709 or sRGB. However the eye can see the difference between the colour levels, especially when there is a sharp border between two large areas of adjacent colour levels. This will happen with gradual gradients (like sunsets, dawns or clear blue skies), and also when blurring an image a large amount. Colour banding is more noticeable with fewer bits per pixel (BPP) at 16–256 colours (4–8 BPP), where there are fewer shades with a larger difference between them. The appearance of colour banding is exaggerated by the Mach bands effect. Possible solutions include the introduction of dithering and increasing the number of bits per colour channel. Because the banding comes from limitations in the presentation of the image, blurring the image does not fix this unless the image BPP is higher than the original.

Foveated rendering

Foveated rendering is a rendering technique which uses an eye tracker integrated with a virtual reality headset to reduce the rendering workload by greatly reducing the image quality in the peripheral vision (outside of the zone gazed by the fovea). A less sophisticated variant called fixed foveated rendering doesn't utilise eye tracking and instead assumes a fixed focal point. == History == Research into foveated rendering dates back at least to 1991. At Tech Crunch Disrupt SF 2014, Fove unveiled a headset featuring foveated rendering. This was followed by a successful kickstarter in May 2015. At CES 2016, SensoMotoric Instruments (SMI) demoed a new 250 Hz eye tracking system and a working foveated rendering solution. It resulted from a partnership with camera sensor manufacturer Omnivision who provided the camera hardware for the new system. In July 2016, Nvidia demonstrated during SIGGRAPH a new method of foveated rendering claimed to be invisible to users. In February 2017, Qualcomm announced their Snapdragon 835 Virtual Reality Development Kit (VRDK) which includes foveated rendering support called Adreno Foveation. == Use == According to chief scientist Michael Abrash at Oculus, utilising foveated rendering in conjunction with sparse rendering and deep learning image reconstruction has the potential to require an order of magnitude fewer pixels to be rendered in comparison to a full image. Later, these results have been demonstrated and published. In December 2019, fixed foveated rendering support was added to the Oculus Quest SDK. A number of VR headsets have included on-board eye tracking to provide support for foveated rendering, including HTC's Vive Pro Eye (2019), Meta Quest Pro (2022), PlayStation VR2 (2023), and Apple Vision Pro (2024). In 2025, Valve announced the upcoming Steam Frame headset, which applies a variation of the technique known as "foveated streaming" for wireless streaming from a PC to the headset; the method similarly uses variance in bit rate, and is performed at the encoder level rather than the software level.

Genetic operator

A genetic operator is an operator used in evolutionary algorithms (EA) to guide the algorithm towards a solution to a given problem. There are three main types of operators (mutation, crossover and selection), which must work in conjunction with one another in order for the algorithm to be successful. Genetic operators are used to create and maintain genetic diversity (mutation operator), combine existing solutions (also known as chromosomes) into new solutions (crossover) and select between solutions (selection). The classic representatives of evolutionary algorithms include genetic algorithms, evolution strategies, genetic programming and evolutionary programming. In his book discussing the use of genetic programming for the optimization of complex problems, computer scientist John Koza has also identified an 'inversion' or 'permutation' operator; however, the effectiveness of this operator has never been conclusively demonstrated and this operator is rarely discussed in the field of genetic programming. For combinatorial problems, however, these and other operators tailored to permutations are frequently used by other EAs. Mutation (or mutation-like) operators are said to be unary operators, as they only operate on one chromosome at a time. In contrast, crossover operators are said to be binary operators, as they operate on two chromosomes at a time, combining two existing chromosomes into one new chromosome. == Operators == Genetic variation is a necessity for the process of evolution. Genetic operators used in evolutionary algorithms are analogous to those in the natural world: survival of the fittest, or selection; reproduction (crossover, also called recombination); and mutation. === Selection === Selection operators give preference to better candidate solutions (chromosomes), allowing them to pass on their 'genes' to the next generation (iteration) of the algorithm. The best solutions are determined using some form of objective function (also known as a 'fitness function' in evolutionary algorithms), before being passed to the crossover operator. Different methods for choosing the best solutions exist, for example, fitness proportionate selection and tournament selection. A further or the same selection operator is used to determine the individuals for being selected to form the next parental generation. The selection operator may also ensure that the best solution(s) from the current generation always become(s) a member of the next generation without being altered; this is known as elitism or elitist selection. === Crossover === Crossover is the process of taking more than one parent solutions (chromosomes) and producing a child solution from them. By recombining portions of good solutions, the evolutionary algorithm is more likely to create a better solution. As with selection, there are a number of different methods for combining the parent solutions, including the edge recombination operator (ERO) and the 'cut and splice crossover' and 'uniform crossover' methods. The crossover method is often chosen to closely match the chromosome's representation of the solution; this may become particularly important when variables are grouped together as building blocks, which might be disrupted by a non-respectful crossover operator. Similarly, crossover methods may be particularly suited to certain problems; the ERO is considered a good option for solving the travelling salesman problem. === Mutation === The mutation operator encourages genetic diversity amongst solutions and attempts to prevent the evolutionary algorithm converging to a local minimum by stopping the solutions becoming too close to one another. In mutating the current pool of solutions, a given solution may change between slightly and entirely from the previous solution. By mutating the solutions, an evolutionary algorithm can reach an improved solution solely through the mutation operator. Again, different methods of mutation may be used; these range from a simple bit mutation (flipping random bits in a binary string chromosome with some low probability) to more complex mutation methods in which genes in the solution are changed, for example by adding a random value from the Gaussian distribution to the current gene value. As with the crossover operator, the mutation method is usually chosen to match the representation of the solution within the chromosome. == Combining operators == While each operator acts to improve the solutions produced by the evolutionary algorithm working individually, the operators must work in conjunction with each other for the algorithm to be successful in finding a good solution. Using the selection operator on its own will tend to fill the solution population with copies of the best solution from the population. If the selection and crossover operators are used without the mutation operator, the algorithm will tend to converge to a local minimum, that is, a good but sub-optimal solution to the problem. Using the mutation operator on its own leads to a random walk through the search space. Only by using all three operators together can the evolutionary algorithm become a noise-tolerant global search algorithm, yielding good solutions to the problem at hand.

Mixture model

In statistics, a mixture model is a probabilistic model for representing the presence of subpopulations within an overall population, without requiring that an observed data set should identify the sub-population to which an individual observation belongs. Formally a mixture model corresponds to the mixture distribution that represents the probability distribution of observations in the overall population. However, while problems associated with "mixture distributions" relate to deriving the properties of the overall population from those of the sub-populations, "mixture models" are used to make statistical inferences about the properties of the sub-populations given only observations on the pooled population, without sub-population identity information. Mixture models are used for clustering, under the name model-based clustering, and also for density estimation. Mixture models should not be confused with models for compositional data, i.e., data whose components are constrained to sum to a constant value (1, 100%, etc.). However, compositional models can be thought of as mixture models, where members of the population are sampled at random. Conversely, mixture models can be thought of as compositional models, where the total size reading population has been normalized to 1. == Structure == === General mixture model === A typical finite-dimensional mixture model is a hierarchical model consisting of the following components: N random variables that are observed, each distributed according to a mixture of K components, with the components belonging to the same parametric family of distributions (e.g., all normal, all Zipfian, etc.) but with different parameters. However, it is also possible to have a finite mixture model where each component belongs to a different parametric family of distributions, for example, a mixture of a multivariate normal distribution and a generalized hyperbolic distribution. N random latent variables specifying the identity of the mixture component of each observation, each distributed according to a K-dimensional categorical distribution A set of K mixture weights, which are probabilities that sum to 1. A set of K parameters, each specifying the parameter of the corresponding mixture component. In many cases, each "parameter" is actually a set of parameters. For example, if the mixture components are Gaussian distributions, there will be a mean and variance for each component. If the mixture components are categorical distributions (e.g., when each observation is a token from a finite alphabet of size V), there will be a vector of V probabilities summing to 1. In addition, in a Bayesian setting, the mixture weights and parameters will themselves be random variables, and prior distributions will be placed over the variables. In such a case, the weights are typically viewed as a K-dimensional random vector drawn from a Dirichlet distribution (the conjugate prior of the categorical distribution), and the parameters will be distributed according to their respective conjugate priors. Mathematically, a basic parametric mixture model can be described as follows: K = number of mixture components N = number of observations θ i = 1 … K = parameter of distribution of observation associated with component i ϕ i = 1 … K = mixture weight, i.e., prior probability of a particular component i ϕ = K -dimensional vector composed of all the individual ϕ 1 … K ; must sum to 1 z i = 1 … N = component of observation i x i = 1 … N = observation i F ( x | θ ) = probability distribution of an observation, parametrized on θ z i = 1 … N ∼ Categorical ⁡ ( ϕ ) x i = 1 … N | z i = 1 … N ∼ F ( θ z i ) {\displaystyle {\begin{array}{lcl}K&=&{\text{number of mixture components}}\\N&=&{\text{number of observations}}\\\theta _{i=1\dots K}&=&{\text{parameter of distribution of observation associated with component }}i\\\phi _{i=1\dots K}&=&{\text{mixture weight, i.e., prior probability of a particular component }}i\\{\boldsymbol {\phi }}&=&K{\text{-dimensional vector composed of all the individual }}\phi _{1\dots K}{\text{; must sum to 1}}\\z_{i=1\dots N}&=&{\text{component of observation }}i\\x_{i=1\dots N}&=&{\text{observation }}i\\F(x|\theta )&=&{\text{probability distribution of an observation, parametrized on }}\theta \\z_{i=1\dots N}&\sim &\operatorname {Categorical} ({\boldsymbol {\phi }})\\x_{i=1\dots N}|z_{i=1\dots N}&\sim &F(\theta _{z_{i}})\end{array}}} In a Bayesian setting, all parameters are associated with random variables, as follows: K , N = as above θ i = 1 … K , ϕ i = 1 … K , ϕ = as above z i = 1 … N , x i = 1 … N , F ( x | θ ) = as above α = shared hyperparameter for component parameters β = shared hyperparameter for mixture weights H ( θ | α ) = prior probability distribution of component parameters, parametrized on α θ i = 1 … K ∼ H ( θ | α ) ϕ ∼ S y m m e t r i c - D i r i c h l e t K ⁡ ( β ) z i = 1 … N | ϕ ∼ Categorical ⁡ ( ϕ ) x i = 1 … N | z i = 1 … N , θ i = 1 … K ∼ F ( θ z i ) {\displaystyle {\begin{array}{lcl}K,N&=&{\text{as above}}\\\theta _{i=1\dots K},\phi _{i=1\dots K},{\boldsymbol {\phi }}&=&{\text{as above}}\\z_{i=1\dots N},x_{i=1\dots N},F(x|\theta )&=&{\text{as above}}\\\alpha &=&{\text{shared hyperparameter for component parameters}}\\\beta &=&{\text{shared hyperparameter for mixture weights}}\\H(\theta |\alpha )&=&{\text{prior probability distribution of component parameters, parametrized on }}\alpha \\\theta _{i=1\dots K}&\sim &H(\theta |\alpha )\\{\boldsymbol {\phi }}&\sim &\operatorname {Symmetric-Dirichlet} _{K}(\beta )\\z_{i=1\dots N}|{\boldsymbol {\phi }}&\sim &\operatorname {Categorical} ({\boldsymbol {\phi }})\\x_{i=1\dots N}|z_{i=1\dots N},\theta _{i=1\dots K}&\sim &F(\theta _{z_{i}})\end{array}}} This characterization uses F and H to describe arbitrary distributions over observations and parameters, respectively. Typically H will be the conjugate prior of F. The two most common choices of F are Gaussian aka "normal" (for real-valued observations) and categorical (for discrete observations). Other common possibilities for the distribution of the mixture components are: Binomial distribution, for the number of "positive occurrences" (e.g., successes, yes votes, etc.) given a fixed number of total occurrences Multinomial distribution, similar to the binomial distribution, but for counts of multi-way occurrences (e.g., yes/no/maybe in a survey) Negative binomial distribution, for binomial-type observations but where the quantity of interest is the number of failures before a given number of successes occurs Poisson distribution, for the number of occurrences of an event in a given period of time, for an event that is characterized by a fixed rate of occurrence Exponential distribution, for the time before the next event occurs, for an event that is characterized by a fixed rate of occurrence Log-normal distribution, for positive real numbers that are assumed to grow exponentially, such as incomes or prices Multivariate normal distribution (aka multivariate Gaussian distribution), for vectors of correlated outcomes that are individually Gaussian-distributed Multivariate Student's t-distribution, for vectors of heavy-tailed correlated outcomes A vector of Bernoulli-distributed values, corresponding, e.g., to a black-and-white image, with each value representing a pixel; see the handwriting-recognition example below === Specific examples === ==== Gaussian mixture model ==== A typical non-Bayesian Gaussian mixture model looks like this: K , N = as above ϕ i = 1 … K , ϕ = as above z i = 1 … N , x i = 1 … N = as above θ i = 1 … K = { μ i = 1 … K , σ i = 1 … K 2 } μ i = 1 … K = mean of component i σ i = 1 … K 2 = variance of component i z i = 1 … N ∼ Categorical ⁡ ( ϕ ) x i = 1 … N ∼ N ( μ z i , σ z i 2 ) {\displaystyle {\begin{array}{lcl}K,N&=&{\text{as above}}\\\phi _{i=1\dots K},{\boldsymbol {\phi }}&=&{\text{as above}}\\z_{i=1\dots N},x_{i=1\dots N}&=&{\text{as above}}\\\theta _{i=1\dots K}&=&\{\mu _{i=1\dots K},\sigma _{i=1\dots K}^{2}\}\\\mu _{i=1\dots K}&=&{\text{mean of component }}i\\\sigma _{i=1\dots K}^{2}&=&{\text{variance of component }}i\\z_{i=1\dots N}&\sim &\operatorname {Categorical} ({\boldsymbol {\phi }})\\x_{i=1\dots N}&\sim &{\mathcal {N}}(\mu _{z_{i}},\sigma _{z_{i}}^{2})\end{array}}} A Bayesian version of a Gaussian mixture model is as follows: K , N = as above ϕ i = 1 … K , ϕ = as above z i = 1 … N , x i = 1 … N = as above θ i = 1 … K = { μ i = 1 … K , σ i = 1 … K 2 } μ i = 1 … K = mean of component i σ i = 1 … K 2 = variance of component i μ 0 , λ , ν , σ 0 2 = shared hyperparameters μ i = 1 … K ∼ N ( μ 0 , λ σ i 2 ) σ i = 1 … K 2 ∼ I n v e r s e - G a m m a ⁡ ( ν , σ 0 2 ) ϕ ∼ S y m m e t r i c - D i r i c h l e t K ⁡ ( β ) z i = 1 … N ∼ Categorical ⁡ ( ϕ ) x i = 1 … N ∼ N ( μ z i , σ z i 2 ) {\displaystyle {\begin{array}{lcl}K,N&=&{\text{as above}}\\\phi _{i=1\dots K},{\boldsymbol {\phi }}&=&{\text{as above}}\\z_{i=1\dots N},x_{i=1\dots N}&=&{\text{as above}}\\\theta _{i=1\

Ensemble learning

In statistics and machine learning, ensemble methods use multiple learning algorithms to obtain better predictive performance than could be obtained from any of the constituent learning algorithms alone. Unlike a statistical ensemble in statistical mechanics, which is usually infinite, a machine learning ensemble consists of only a concrete finite set of alternative models, but typically allows for much more flexible structure to exist among those alternatives. == Overview == Supervised learning algorithms search through a hypothesis space to find a suitable hypothesis that will make good predictions with a particular problem. Even if this space contains hypotheses that are very well-suited for a particular problem, it may be very difficult to find a good one. Ensembles combine multiple hypotheses to form one which should be theoretically better. Ensemble learning trains two or more machine learning algorithms on a specific classification or regression task. The algorithms within the ensemble model are generally referred as "base models", "base learners", or "weak learners" in literature. These base models can be constructed using a single modelling algorithm, or several different algorithms. The idea is to train a diverse set of weak models on the same modelling task, such that the outputs of each weak learner have poor predictive ability (i.e., high bias), and among all weak learners, the outcome and error values exhibit high variance. Fundamentally, an ensemble learning model trains at least two high-bias (weak) and high-variance (diverse) models to be combined into a better-performing model. The set of weak models — which would not produce satisfactory predictive results individually — are combined or averaged to produce a single, high performing, accurate, and low-variance model to fit the task as required. Ensemble learning typically refers to bagging (bootstrap aggregating), boosting or stacking/blending techniques to induce high variance among the base models. Bagging creates diversity by generating random samples from the training observations and fitting the same model to each different sample — also known as homogeneous parallel ensembles. Boosting follows an iterative process by sequentially training each base model on the up-weighted errors of the previous base model, producing an additive model to reduce the final model errors — also known as sequential ensemble learning. Stacking or blending consists of different base models, each trained independently (i.e. diverse/high variance) to be combined into the ensemble model — producing a heterogeneous parallel ensemble. Common applications of ensemble learning include random forests (an extension of bagging), Boosted Tree models, and Gradient Boosted Tree Models. Models in applications of stacking are generally more task-specific — such as combining clustering techniques with other parametric and/or non-parametric techniques. Evaluating the prediction of an ensemble typically requires more computation than evaluating the prediction of a single model. In one sense, ensemble learning may be thought of as a way to compensate for poor learning algorithms by performing a lot of extra computation. On the other hand, the alternative is to do a lot more learning with one non-ensemble model. An ensemble may be more efficient at improving overall accuracy for the same increase in compute, storage, or communication resources by using that increase on two or more methods, than would have been improved by increasing resource use for a single method. Fast algorithms such as decision trees are commonly used in ensemble methods (e.g., random forests), although slower algorithms can benefit from ensemble techniques as well. By analogy, ensemble techniques have been used also in unsupervised learning scenarios, for example in consensus clustering or in anomaly detection. == Ensemble theory == Empirically, ensembles tend to yield better results when there is a significant diversity among the models. Many ensemble methods, therefore, seek to promote diversity among the models they combine. Although perhaps non-intuitive, more random algorithms (like random decision trees) can be used to produce a stronger ensemble than very deliberate algorithms (like entropy-reducing decision trees). Using a variety of strong learning algorithms, however, has been shown to be more effective than using techniques that attempt to dumb-down the models in order to promote diversity. It is possible to increase diversity in the training stage of the model using correlation for regression tasks or using information measures such as cross entropy for classification tasks. Theoretically, one can justify the diversity concept because the lower bound of the error rate of an ensemble system can be decomposed into accuracy, diversity, and the other term. === The geometric framework === Ensemble learning, including both regression and classification tasks, can be explained using a geometric framework. Within this framework, the output of each individual classifier or regressor for the entire dataset can be viewed as a point in a multi-dimensional space. Additionally, the target result is also represented as a point in this space, referred to as the "ideal point." The Euclidean distance is used as the metric to measure both the performance of a single classifier or regressor (the distance between its point and the ideal point) and the dissimilarity between two classifiers or regressors (the distance between their respective points). This perspective transforms ensemble learning into a deterministic problem. For example, within this geometric framework, it can be proved that the averaging of the outputs (scores) of all base classifiers or regressors can lead to equal or better results than the average of all the individual models. It can also be proved that if the optimal weighting scheme is used, then a weighted averaging approach can outperform any of the individual classifiers or regressors that make up the ensemble or as good as the best performer at least. == Ensemble size == While the number of component classifiers of an ensemble has a great impact on the accuracy of prediction, there is a limited number of studies addressing this problem. A priori determining of ensemble size and the volume and velocity of big data streams make this even more crucial for online ensemble classifiers. Mostly statistical tests were used for determining the proper number of components. More recently, a theoretical framework suggested that there is an ideal number of component classifiers for an ensemble such that having more or less than this number of classifiers would deteriorate the accuracy. It is called "the law of diminishing returns in ensemble construction." Their theoretical framework shows that using the same number of independent component classifiers as class labels gives the highest accuracy. == Common types of ensembles == === Bayes optimal classifier === The Bayes optimal classifier is a classification technique. It is an ensemble of all the hypotheses in the hypothesis space. On average, no other ensemble can outperform it. The Naive Bayes classifier is a version of this that assumes that the data is conditionally independent on the class and makes the computation more feasible. Each hypothesis is given a vote proportional to the likelihood that the training dataset would be sampled from a system if that hypothesis were true. To facilitate training data of finite size, the vote of each hypothesis is also multiplied by the prior probability of that hypothesis. The Bayes optimal classifier can be expressed with the following equation: y = a r g m a x c j ∈ C ∑ h i ∈ H P ( c j | h i ) P ( T | h i ) P ( h i ) {\displaystyle y={\underset {c_{j}\in C}{\mathrm {argmax} }}\sum _{h_{i}\in H}{P(c_{j}|h_{i})P(T|h_{i})P(h_{i})}} where y {\displaystyle y} is the predicted class, C {\displaystyle C} is the set of all possible classes, H {\displaystyle H} is the hypothesis space, P {\displaystyle P} refers to a probability, and T {\displaystyle T} is the training data. As an ensemble, the Bayes optimal classifier represents a hypothesis that is not necessarily in H {\displaystyle H} . The hypothesis represented by the Bayes optimal classifier, however, is the optimal hypothesis in ensemble space (the space of all possible ensembles consisting only of hypotheses in H {\displaystyle H} ). This formula can be restated using Bayes' theorem, which says that the posterior is proportional to the likelihood times the prior: P ( h i | T ) ∝ P ( T | h i ) P ( h i ) {\displaystyle P(h_{i}|T)\propto P(T|h_{i})P(h_{i})} hence, y = a r g m a x c j ∈ C ∑ h i ∈ H P ( c j | h i ) P ( h i | T ) {\displaystyle y={\underset {c_{j}\in C}{\mathrm {argmax} }}\sum _{h_{i}\in H}{P(c_{j}|h_{i})P(h_{i}|T)}} === Bootstrap aggregating (bagging) === Bootstrap aggregation (bagging) involves training an ensemble on bootstrapped data sets. A bootstrapped set is cr

The Outliner of Giants

The Outliner of Giants was commercial outlining software. Like other outliners, it allowed the user to create a document consisting of a series of nested lists. It was one of a number of browser-based outliners that are delivered as a web application, used through a web browser, rather than being installed as a stand-alone application. The Outliner of Giants was released in 2009. The service was shut down on December 31, 2017 and only exports are allowed at this time. == Feature set == Unlike most other browser-based outliners - which often focus on providing a minimum viable product - the Outliner of Giants had much of the functionality typically associated with a desktop outliner, such as the ability to use of columns to structure information. However, The Outliner of Giants did not support offline editing, requiring an active internet connection in order to make changes to an outline document. === Outlining === Like all outliners, The Outliner of Giants supported the creation of a hierarchy of items, with users modifying the parent-child relationship between items in order to structure a document. This included the ability to promote or demote items up or down the hierarchy, or move an item up or down a list of siblings on the same level. The Outliner of Giants did not support the true cloning of items (where an item can appear to be in multiple places within the hierarchy at the same time), although it did support the copying of single or multiple nodes. === Import === The Outliner of Giants could import both plain text and the OPML XML format, which is commonly used to transfer data between outlining applications. === Editing === Outline documents could be edited using a WYSIWYG editor, as well as the Markdown, and Textile markup languages. === Annotation === The Outliner of Giants supported functions to annotate an outline, such as the ability to add colored labels, highlights and text, as well as tags and hashtags. === Collaboration === The Outliner of Giants supported real-time collaboration, where multiple users could edit the same document, and can see the changes made by another user as they happened. === Publication === Outlines created through The Outliner of Giants could be published directly online through the service, either as outlines, pages or in a blog format. === Export === The Outliner of Giants can export outline data as plain text, HTML, as well as directly to the Google Docs word processor.

Canonical correspondence analysis

In multivariate analysis, canonical correspondence analysis (CCA) is an ordination technique that determines axes from the response data as a unimodal combination of measured predictors. CCA is commonly used in ecology in order to extract gradients that drive the composition of ecological communities. CCA extends correspondence analysis (CA) with regression, in order to incorporate predictor variables. == History == CCA was developed in 1986 by Cajo ter Braak and implemented in the program CANOCO, an extension of DECORANA. To date, CCA is one of the most popular multivariate methods in ecology, despite the availability of contemporary alternatives. CCA was originally derived and implemented using an algorithm of weighted averaging, though Legendre & Legendre (1998) derived an alternative algorithm. == Assumptions == The requirements of a CCA are that the samples are random and independent. Also, the data are categorical and that the independent variables are consistent within the sample site and error-free. The original publication states the need for equal species tolerances, equal species maxima, and equispaced or uniformly distributed species optima and site scores.